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On The Cover: The photosynthetic water-oxidizing Mn₄Ca cluster in photosystem II is located on the luminal side of the thylakoid membrane of chloroplasts in the leafs of plants. The details about the orientation and structure of the Mn₄Ca cluster (manganese is red, calcium is green, and oxygen is brown) in the membrane are described in the article by Pushkar et al., pages 7198–7208.

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